Another tendency is to parallel trails more than walking on the more established trails. You may have a parade of deer, including 2 1/2 & 3 1/2 yr old bucks come by. Later, the mature one RUNS by, parallel to the others (typically downwind of the trail).
Which brings up another trait:
They have more a tendency to run from Point A to Point B,
as opposed to simply slipping along a trail.
The mature one may stand motionless for a long period as the other deer mosey along. Then suddenly, he just takes off in a flat out run, only to stop again, and stand for a long period.
Making them even more unpredictable, they are more likely to "cut a diagonal", and do so in a flat out run.
They may be following along behind other deer, off-set to the other deer's trail by 75 yds, perhaps 150 yds behind them, but the deer they're "following" are on a trail that twists & curves. Next thing you know, the mature buck takes a "short cut" by "cutting the diagonal", maybe only needing to periodically "run" 150 yds for every 250 yds the other deer are more steadily just mosying along.