The 'lands' are the beginning of the rifling. They are tapered at the chamber before they get to full height- this tapering is known as the 'leade'. Generally rifles with a shorter leade and rounds that are loaded so that the bullet just touches the lands when seated are more accurate. However pressure is increased. A lot of high intensity rounds such as the Weatherbys have long leades to help reduce pressure. Sometimes due to magazine limitations bullets can't be seated long enough to 'kiss' the lands. And some manufacturers like Barnes with the solid copper bullets recommend seating .0010 or so off the lands to increase accuracy(and reduce pressure). Every rifle seems to be pretty much an individual so experimentation with seating depth can determine the sweet spot for seating depth. Of course all of this is influenced by bullet weight(length) and the shape of the ogive too. Personally I don't get too excited about all this- I just load to standard oal paying attention to mag length and go hunting. This works for me with the type of shooting and hunting I do. PH